My problem was that whenever I sent an email from my boogly.net email (hosted by hostgator) to my email at celiacclub.com (email managed with Google Apps for domains), the message bounced, replying with this error:

550-Verification failed for <email>

550-No Such User Here

550 Sender verify failed (state 14).

It only happened for the boogly.net-to/from-celiacclub.com connection–emails from hotmail got through successfully.  If your email isn’t working at all for communication with any email, refer to the Google Configuring MX records page and use dnstools.com to check your work.  Also, make sure your SPF record is set correctly.  At first I thought it was an issue because I’m running Google Apps for Domains.  Thought something was misconfigured on my DNS.  But it wasn’t.  It took a while before I found the Gmail Troubleshooter.  This told me that it was for sure the other host that was attempting to check that the user exists on Google Apps (This is supposed to help alleviate junk mail).

After exchanging emails with hostgator support, I found that the issue was with the hostgator servers’ configuration of localdomains/remotedomains.  In other words, boogly.net thought that celiacclub.com was a local domain so it tried to look on its own system to validate that email address.  You’ll need to modify your server’s awareness of the site.  Here is a good tutorial about how to go about changing this.  Of course, you’ll need to move from the localdomains to the remotedomains.  Depending on your host, you might not be able to modify these files–in which case, you’ll need to get on the phone with their support and have them do this process.

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